![\"\"](\"https:\/\/dev.academist-cf.com\/journal\/wp-content\/uploads\/2017\/07\/001-8.jpg\")
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\u30ec\u30d7\u30ea\u30bd\u30fc\u30e0\u306e\u69cb\u9020\u306f\u67d4\u8edf\u306a\u3082\u306e\u3067\u3042\u308a\u3001\u30d8\u30ea\u30ab\u30fc\u30bc\u306b\u3088\u308b\u8907\u88fd\u30d5\u30a9\u30fc\u30af\u306e\u9032\u884c\u3068DNA\u5408\u6210\u304c\u4e56\u96e2\u3057\u3066\u3057\u307e\u3046\u3053\u3068\u3082\u3042\u308a\u307e\u3059\u3002\u7279\u306b\u3001DNA\u306e\u69cb\u9020\u304c\u640d\u50b7\u3057\u305f\u5834\u5408\u306b\u306f\u3001DNA\u5408\u6210\u306e\u53cd\u5fdc\u304c\u505c\u6b62\u3057\u3001\u305d\u306e\u7d50\u679c\u3068\u3057\u3066\u3001DNA\u640d\u50b7\u306e\u90e8\u5206\u3092\u92f3\u578b\u3068\u3059\u308b\u5408\u6210\u3092\u5f8c\u56de\u3057\u306b\u3057\u3064\u3064\u3001DNA\u8907\u88fd\u304c\u884c\u308f\u308c\u308b\u3053\u3068\u304c\u3042\u308a\u307e\u3059\u3002\u307e\u305f\u3001\u30d2\u30c8\u7d30\u80de\u3092\u542b\u3080\u771f\u6838\u751f\u7269\u306e\u30ec\u30d7\u30ea\u30bd\u30fc\u30e0\u306f\u3001\u5927\u8178\u83cc\u306a\u3069\u306e\u7d30\u83cc\u306e\u3082\u306e\u307b\u3069\u5f37\u56fa\u306a\u3082\u306e\u3067\u306f\u306a\u3044\u3068\u8003\u3048\u3089\u308c\u3066\u3044\u307e\u3059\u3002\u305d\u306e\u3088\u3046\u306a\u610f\u5473\u3067\u3001DNA\u30dd\u30ea\u30e1\u30e9\u30fc\u30bc\u306b\u3088\u308b\u5408\u6210\u53cd\u5fdc\u304cDNA\u8907\u88fd\u30d5\u30a9\u30fc\u30af\u306e\u5f8c\u65b9\u3067\u81ea\u5f8b\u7684\u306b\u6a5f\u80fd\u3059\u308b\u3053\u3068\u304c\u5fc5\u8981\u3067\u3042\u308a\u3001\u305d\u308c\u3092\u5236\u5fa1\u3059\u308b\u305f\u3081\u306b\u3001\u300c\u8907\u88fd\u30af\u30e9\u30f3\u30d7\u300d\u304c\u91cd\u8981\u306a\u5f79\u5272\u3092\u679c\u305f\u3057\u307e\u3059\u3002<\/p>\n
![\"\"](\"https:\/\/dev.academist-cf.com\/journal\/wp-content\/uploads\/2017\/07\/002-8.jpg\")
<\/p>\n
\u30c9\u30fc\u30ca\u30c4\u72b6\u5206\u5b50DNA\u8907\u88fd\u30af\u30e9\u30f3\u30d7\u304cDNA\u5408\u6210\u306e\u73fe\u5834\u76e3\u7763\u3068\u3057\u3066\u50cd\u304f<\/h3>\n
DNA\u8907\u88fd\u306b\u5fc5\u8981\u306aDNA\u30dd\u30ea\u30e1\u30e9\u30fc\u30bc\u306a\u3069\u306e\u8907\u88fd\u306b\u95a2\u308f\u308b\u9175\u7d20\u306e\u591a\u304f\u306f\u3001\u8907\u88fd\u30af\u30e9\u30f3\u30d7\u3068\u3044\u3046\u5206\u5b50\u3092\u4ecb\u3057\u3066\u3001DNA\u3068\u76f8\u4e92\u4f5c\u7528\u3092\u3057\u307e\u3059\u3002\u8907\u88fd\u30af\u30e9\u30f3\u30d7\u306f\u30c9\u30fc\u30ca\u30c3\u30c4\u72b6\u306e\u69cb\u9020\u3092\u3068\u308a\u3001RFC\u3068\u547c\u3070\u308c\u308b\u30af\u30e9\u30f3\u30d7\u30ed\u30fc\u30c0\u30fc\u5206\u5b50\u306b\u3088\u3063\u3066\u3001\u751f\u4f53\u30a8\u30cd\u30eb\u30ae\u30fc\u4f9d\u5b58\u7684\u306b\u305d\u306e\u8f2a\u304c\u958b\u304b\u308c\u3001DNA\u4e0a\u306b\u8f09\u305b\u3089\u308c\u307e\u3059\u3002DNA\u4e0a\u306e\u30af\u30e9\u30f3\u30d7DNA\u306f\u7cf8\u306b\u901a\u3057\u305f\u8f2a\u306e\u3088\u3046\u306b\u79fb\u52d5\u53ef\u80fd\u3067\u3042\u308a\u3001\u8907\u88fd\u30af\u30e9\u30f3\u30d7\u3068\u5171\u306bDNA\u3092\u5408\u6210\u3059\u308b\u9175\u7d20\uff08DNA\u30dd\u30ea\u30e1\u30e9\u30fc\u30bc\uff09\u304c\u30b9\u30e9\u30a4\u30c9\u3057\u3001\u30b9\u30e0\u30fc\u30ba\u306aDNA\u5408\u6210\u304c\u884c\u308f\u308c\u307e\u3059\u3002\u3053\u308c\u306f\u3001\u7d30\u83cc\u304b\u3089\u771f\u6838\u751f\u7269\u307e\u3067\u306b\u4fdd\u5b58\u3055\u308c\u308b\u4ed5\u7d44\u307f\u3067\u3059\u3002\u771f\u6838\u751f\u7269\u306e\u5834\u5408\u306f\u3001PCNA\uff08Proliferating Cell Nuclear Antigen\uff09\u3068\u547c\u3070\u308c\u307e\u3059\u3002\u3082\u3068\u3082\u3068\u3001\u3088\u304f\u5897\u6b96\u3059\u308b\u7d30\u80de\u306e\u6838\u306b\u591a\u304f\u5b58\u5728\u3059\u308b\u30bf\u30f3\u30d1\u30af\u8cea\u3068\u3057\u3066\u767a\u898b\u3055\u308c\u305f\u305f\u3081\u3001\u3053\u306e\u3088\u3046\u306a\u540d\u524d\u306b\u306a\u3063\u3066\u3044\u307e\u3059\u3002<\/p>\n
PCNA\u306f\u5358\u306bDNA\u30dd\u30ea\u30e1\u30e9\u30fc\u30bc\u306a\u3069\u306e\u9175\u7d20\u306e\u50cd\u304d\u3092\u52a9\u3051\u308b\u3060\u3051\u3067\u306a\u304f\u3001DNA\u5408\u6210\u304c\u6ede\u3063\u305f\u3068\u304d\u306a\u3069\u306b\u3001\u5fc5\u8981\u306a\u30bf\u30f3\u30d1\u30af\u8cea\u3092\u305d\u306e\u5834\u306b\u547c\u3073\u8fbc\u3080\u5f79\u5272\u3082\u3042\u308a\u307e\u3059\u3002DNA\u5408\u6210\u304c\u6ede\u3063\u305f\u5834\u5408\u306f\u5fc5\u7136\u7684\u306b\u3001DNA\u30dd\u30ea\u30e1\u30e9\u30fc\u30bc\u306e\u524d\u65b9\u306b\u92f3\u578b\u3068\u306a\u308b\u77ed\u9396DNA\u304c\u3080\u304d\u51fa\u3057\u306b\u306a\u308a\u3001\u305d\u308c\u304c\u304d\u3063\u304b\u3051\u3067\u3001PCNA\u304c\u30e6\u30d3\u30ad\u30c1\u30f3\u3068\u3044\u3046\u5c0f\u3055\u306a\u5206\u5b50\u306e\u4fee\u98fe\u3092\u53d7\u3051\u307e\u3059\u3002\u30e6\u30d3\u30ad\u30c1\u30f3\u5316\u306f\u672c\u6765\u3001\u30bf\u30f3\u30d1\u30af\u8cea\u3092\u5206\u89e3\u3059\u308b\u305f\u3081\u306e\u6307\u6a19\u3068\u306a\u308b\u73fe\u8c61\u3067\u3059\u304c\u3001\u3053\u306e\u5834\u5408\u306f\u3001\u30e6\u30d3\u30ad\u30c1\u30f3\u5316\u3057\u305fPCNA\u304cDNA\u306e\u640d\u50b7\u90e8\u4f4d\u306b\u7559\u307e\u308a\u3001DNA\u640d\u50b7\u3092\u4e57\u308a\u8d8a\u3048\u3066\u5408\u6210\u3092\u884c\u3048\u308bDNA\u30dd\u30ea\u30e1\u30e9\u30fc\u30bc\u306e\u8db3\u5834\u3068\u306a\u308a\u307e\u3059\uff08\u4ee5\u4e0b\u306e\u56f3\u4e2d\uff08i\uff09\uff09\u3002<\/p>\n
\u6211\u3005\u306e\u7814\u7a76\u30b0\u30eb\u30fc\u30d7\u306f\u3001DNA\u640d\u50b7\u306b\u5bfe\u51e6\u3059\u308b\u4ed5\u7d44\u307f\u306b\u52a0\u3048\u3066\u3001PCNA\u306e\u30e6\u30d3\u30ad\u30c1\u30f3\u5316\u304c\u5ca1\u5d0e\u30d5\u30e9\u30b0\u30e1\u30f3\u30c8\u306e\u5408\u6210\u3092\u88dc\u52a9\u3059\u308b\u4ed5\u7d44\u307f\u3067\u3042\u308b\u3053\u3068\u3092\u63d0\u5531\u3057\u307e\u3057\u305f\uff08\u4ee5\u4e0b\u306e\u56f3\u4e2d\uff08ii\uff09\uff09\u3002\u307e\u305a\u3001\u300cPCNA\u306e\u30e6\u30d3\u30ad\u30c1\u30f3\u5316\u304cDNA\u5408\u6210\u3092\u884c\u3063\u3066\u3044\u308b\u7d30\u80de\u3067\u81ea\u7136\u3068\u8d77\u304d\u308b\u73fe\u8c61\u3067\u3042\u308a\u3001DNA\u5408\u6210\u306e\u52b9\u7387\u7684\u306a\u5b8c\u4e86\u306b\u95a2\u308f\u308b\u30d5\u30a1\u30af\u30bf\u30fc\u3067\u3042\u308b\u3053\u3068\u300d\u3092\u793a\u3057\u307e\u3057\u305f\u3002\u307e\u305f\u3001\u30e6\u30d3\u30ad\u30c1\u30f3\u5316PCNA\u306eDNA\u8907\u88fd\u30b7\u30b9\u30c6\u30e0\u306b\u304a\u3051\u308b\u5f79\u5272\u3092\u89e3\u6790\u3057\u305f\u7d50\u679c\u3001PCNA\u304c\u30e6\u30d3\u30ad\u30c1\u30f3\u5316\u306b\u3088\u308aDNA\u9396\u4e0a\u306b\u7559\u307e\u308a\u3084\u3059\u304f\u306a\u308b\u3053\u3068\u3092\u793a\u3057\u3001\u305d\u306e\u7d50\u679c\u3001PCNA\u3068\u30e9\u30ae\u30f3\u30b0\u9396\u5408\u6210\u3092\u304a\u3053\u306a\u3046\u30dd\u30ea\u30e1\u30e9\u30fc\u30bcPol\u03b4\uff08\u30c7\u30eb\u30bf\uff09\u304c\u8907\u88fd\u306e\u5834\u306b\u30a2\u30af\u30bb\u30b9\u3057\u3084\u3059\u304f\u306a\u308b\u3053\u3068\u3092\u793a\u3057\u307e\u3057\u305f\u3002\u30e9\u30ae\u30f3\u30b0\u9396\u5408\u6210\u304c\u6ede\u308a\u77ed\u9396DNA\u306e\u92f3\u578b\u304c\u6b8b\u5b58\u3057\u3066\u3044\u308b\u5834\u5408\u306b\uff0cPCNA\u30e6\u30d3\u30ad\u30c1\u30f3\u5316\u3092\u4ecb\u3057\u3066 Pol\u03b4\u306e\u6a5f\u80fd\u3092\u5897\u5f37\u3057\u3066\u3044\u308b\u3068\u8003\u3048\u3089\u308c\u307e\u3059\u3002\u5ca1\u5d0e\u30d5\u30e9\u30b0\u30e1\u30f3\u30c8\u306f100\u5869\u57fa\u307b\u3069\u77ed\u3044\u9396\u3067\u3042\u308a\u30011\u56de\u306e\u30b2\u30ce\u30e0\u8907\u88fd\u3067\u9175\u6bcd\u306e\u5834\u5408\u306f\u7d04\u5341\u4e07\u56de\u3001\u30d2\u30c8\u306e\u5834\u5408\u306f\u6570\u5343\u4e07\u56de\u3001\u305d\u306e\u5408\u6210\u304c\u7e70\u308a\u8fd4\u3055\u308c\u307e\u3059\u3002\u305d\u306e\u969b\u306b\u8d77\u304d\u3046\u308b\u30a8\u30e9\u30fc\u3092\u9632\u3050\u305f\u3081\u306b\u3001PCNA\u306e\u4fee\u98fe\u306b\u3088\u308bDNA\u5408\u6210\u306e\u73fe\u5834\u3092\u81ea\u5f8b\u7684\u306b\u6307\u63ee\u3059\u308b\u4ed5\u7d44\u307f\u304c\u610f\u7fa9\u3092\u6301\u3064\u306e\u3067\u3057\u3087\u3046\u3002<\/p>\n
![\"\"](\"https:\/\/dev.academist-cf.com\/journal\/wp-content\/uploads\/2017\/07\/003-6.jpg\")
<\/p>\n
\u6700\u5f8c\u306b<\/h3>\n
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<\/p>\n
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\nY. Daigaku, T.J. Etheridge, Y. Nakazawa, M. Nakayama, A.T. Watson, I. Miyabe, T. Ogi, M.A. Osborne, A.M. Carr . PCNA ubiquitylation ensures timely completion of unperturbed DNA replication in fission yeast. PLoS Genet 13(5): e1006789, 2017
\nY. Daigaku, Roadworks of DNA Damage Bypass during and after Replication, Genes and Environment. 34, 77-88, 2012\u3000\uff08\u7dcf\u8aac\uff09
\nY. Daigaku, A. A. Davies and H. D. Ulrich, Ubiquitin-dependent DNA damage bypass is separable from genome replication., Nature. 465, 951-955, 2010<\/p>\n